For our uses, this order could be characterized by the number of species andor the number of individuals in each species, within our sample plot. A measure derived from information theories that were developed by claude elwood shannon 19162001 and norbert wiener. Shannon wiener species diversity index formula data analysis. How to calculate shannon wiener diversity index definition. Natural resource biometrics measures of diversity shannon. A tribute to claude shannon 19162001 and a plea for more. Species diversity species diversity differs from species richness in that it takes into account both the numbers of species present and the dominance or evenness of species in relation to one another. A systematic random sampling approach was adopted to establish 52 sampling plots measuring 10. Prediction of the shannon wiener diversity index from trace element profiles in sediments around the statfjord platforms. Lets compute the shannonweiner diversity index for the same hypothetical community in the previous example. Maximum species evenness was recorded for cyanophyceae and minimum for bacillariophyceae. Shannonwiener index of diversity information index source. Simpson index must also be calculated for this will give you a general idea regarding dominance. Statistics shannon wiener diversity index tutorialspoint.
Shannon wiener index biodiversity studies pdf species diversity of ecological communities is complicated. A tribute to claude shannon 19162001 and a plea for more rigorous use of species richness, species diversity and the shannon wiener index. Diversity, security, and adaptability in energy systems. A greater number of species and a more even distribution both increase diversity as measured by h. The shannon index has been a popular diversity index in the ecological literature, where it is also known as shannons diversity index, the shannonwiener index, the shannonweaver index and the shannon entropy. Proceedings of the 27th workshopconference of the association for biology laboratory education able, 383 pages. However, if you were to include the definitions of those indices, i and others might be able to answer without having to search the literature for those definitions. Species diversity considers both the number of species richness in the community and their relative abundance evenness or equability. Species diversity, abundance and distribution of fish. Diversity or dominance can be calculated with density, cover, or biomass. Type ii indices are most sensitive to changes in the more. The diversity, structure, species composition, and above ground biomass of woody plants on ngomakurira mountain in zimbabwe were studied. The shannon index is a popular diversity index in ecology taking into account the number of species as well as the evenness of species how close in numbers each species in an environment are.
Measurements of biodiversity marinespecies introduced. Diversity indices that combine both richness and abundance i. Simpsons diversity indexis used to calculate a measure of diversity, taking into account the number of something as well as its abundance. This diversity measure is based on information theory. Apr 11, 2003 on some occasions it is called the shannonwiener index and on other occasions it is called the shannonweaver index. Shannon weiner index assumes that individuals are randomly sampled from an independent large population and all the species are represented in the sample. The addition of rare species to a sample causes only small changes in the value of d shannon wiener index also been called the shannon index and the shannon weaver index. Calculate the shannon diversity index and evenness for these sample values. Measuring diversity has been of historical significance and due to the obvious declines in habitat diversity this data is still valuable. The shannon index has been a popular diversity index in.
Many indices for measuring species diversity have been proposed. The shannonwiener diversity index is one measure that we will use to. I have only seen it used for species diversity measure. Posted on 201021 20170904 author bpmsg categories online tools, other articles tags berger parker, calculation, excel, shannon entropy, shannon weaver index, shannonwiener index, simpson index diversity as business kpi alpha and beta diversity.
Dominance indices are weighted toward the abundance of the commonest species. Shannon index shannonweaver index h a diversity index is the measure of species diversity in a given community. B based on the species richness the number of species present and species abundance the number of individuals per species. A diversity index is a mathematical measure of species diversity in a community. The maximum diversity hmax of a sample is found when all species are equally abundant.
The index is most often used for ecological studies that measure species diversity, but the same analysis can also be applied to other principles, such. Sample values s 60,10,25,1,4 number of species n 5. Brief outlines of the two types of diversity indices of biodiversity are discussed in this article. Species and biological diversity choices of diversity. Introduction ondo state is located in south western nigeria and has coastal areas estimated to be about. In this article, a link is noted between a common family of diversity indices and non.
Lets compute the shannon weiner diversity index for the same hypothetical community in the previous example. Distribution, diversity and ecology of spider species at. View enhanced pdf access article on wiley online library html view download pdf for offline viewing. The shannonweiner species diversity index association for. C the more species you have, the more diverse the area.
Shannon weiner index the shannon index is affected by both the number of species and their equitability, or evenness. In particular, for a random sample, we can use shannons index of diversity aka as shannonweiners index, which is. Species diversity concepts describing communities species. Shannon index the idea behind this index is that the diversity of a community is similar to the amount of information in a code or message. Used to compare diversity, doesnt give a measure of dominance. But what else is the shannon index used for michael. Regarding your result, a shannon wiener diversity index of 2. I am not an ecologist and i dont know the shannon wiener index and dont remember by heart the definition of the simpsons index. Diversity of bacterial endophytes in 3 and 15 yearold. A diversity index is the measure of species diversity in a given community. The shannon wiener diversity index is one measure that we will use to try to draw information from samples in the field. What is the correct name for the biodiversity index.
The shannon weiner index is most sensitive to the number of species in a sample, so it is usually considered to be biased toward measuring species richness. We suggest that at the very least, authors should define. Simpson diversity and the shannonwiener index as special. The proportion of species i relative to the total number of species p i is calculated, and then multiplied by the natural logarithm of this proportion lnp i. It is different from species richness in that unlike richness it also shows community composition and takes into account the relative abundance of species that are present in the community. The actual formula for the shannonweiner diversity index is. The shannon weiner diversity index of three different sampling indicated a strong relationship with overall species richness, showed variation and ranged from 0. Aug 28, 2014 in analyses using only one diversity index, similar significant effects of lui would have been detected using any of the indices we included, except e. The most common measures of biodiversity are species richness, simpsons index and shannon wiener index. Simpson diversity and the shannon wiener index as special cases of a generalized entropy. Equitability assumes a value between 0 and 1 with 1 being complete evenness. Pdf application of shannons index to study diversity with. If i obtain the shannonweiner diversity index as 2. It assumed that individuals are randomly sampled from an infinitely large community, and that all species are represented in the sample.
Based on shannon wiener diversity index, the mannwhitney u test showed a significant difference between the two habitats u 2. A a diversity index is a mathematical measure of species diversity in a given community. Shannon wiener index of diversity oxford reference. The shannon index is a measurement used to compare diversity between habitat samples. Application of shannons index to study diversity with. In this study, we also use the shannon wiener index to calculatethe diversity of energy consumption and industry sector. Margalefs diversity index, sorensens index, number of occurrence index noi, shannonwiener index, pielous measure of evenness and berger parker dominance index. Sample values s 60,10,25,1,4 number of species n 5 first, let us calculate the sum of the given values. The shannon index has been a popular diversity index in the ecological. Jan 26, 2012 a diversity index is the measure of species diversity in a given community.
Thus, when conducting simple statistical analyses of a specific effect of disturbance on diversity, the choice of index does not appear particularly important. For categorical data, there is no mean or median and so the measures of variation described in measures of variability cant be used. Key wordsdiversity indices, shannon index, shannon wiener index, shannon weaver index, species richness, species diversity. Diversity index, one of shannon, simpson or invsimpson. In fact, in the late 1940s shannon had built on the work of wiener. This makes the shannon index and the simpson diversity or gini coefficient special cases of a more general index. Key wordsdiversity indices, shannon index, shannon wiener index, shannonweaver index, species richness, species diversity. Shannon index is a commonly used diversity index that takes into account. The actual formula for the shannon weiner diversity index is. Shannon wiener index of diversity published on by oxford university press. This is the simplest of all the measures of species diversity. Pdf on jan 1, 2006, kathleen a nolan and others published beachcomber biology. The shannon weiner diversity index is a common way of showing that diversity involves not only numbers of different species, but also how well each of these species is represented in different habitats.
However, when each representative group was considered as input data, the index resulted 3. This tutorial explains how to calculate the shannon wiener diversity index and evenness. Online biodiversity calculator which helps to calculates the shannon s diversity index and evenness from the given sample data. Instead we use a measure of the distribution of observations among the categories. To do this, the proportion of each species of the total is calculated, the natural log of each is determined, and then multiplied by the proportion. We will then consider one measure used to compare diversity among communities, jaccard. The most widely used diversity index in the ecological literature is the shannon wiener diversity index. In order for an economy to withstand supply and demand shocks, it must either maintain its competitive advantage or have enough variety of industries to reemploy displaced workers malizia and ke, 1993. Values of the shannon diversity index for real communities typically fall between 1.
Application of shannons index to study diversity with reference to. The shannonwiener index is an example of a type i index, like brillouins index. This comparison can be between two different habitats or a comparison of one habitat over time. First, enter the number of species, and then enter the name you wish to give the species, if available, and the given populations for each of the speciesin any given order. Shannon wiener index of diversity information index a measure used by ecologists when a system contains too many individuals for each to be identified and examined. The shannon weiner index was considered to be the most satisfactory measure of energy diversity because it incorporates the concepts of variety and balance 4. Pdf prediction of the shannon wiener diversity index. However, density is the most commonly used variable to assess plant diversity, therefore the examples in this module will be based on density. Jan 06, 2011 demonstration of how to calculate the shannon wiener diversity index in microsoft excel. Species richness, diversity and abundance of some decapod. The shannonweiner species diversity index find, read and cite all the. Diversity indices provide more information about community composition than simply species richness i. Phytoplankton community structure and species diversity of. The shannon diversity index h is another index that is commonly used to characterize species diversity in a community.
Shannons diversity index real statistics using excel. All you do is count of the number of species found in a community e. The highest fish diversity was recorded at siteiii followed by sitesii and i. The shannon weiner value h can range from no diversity at 0. A widely used dominance index is simpsons diversity index.
Here, we will explore two measures of biodiversity. Maximum and minimum species diversity shannon weiner h were found 2. This measure of equitability compares the observed shannonwiener index against. Dpsi log pi, where s is the total number of species in the sample, i is the total number of individuals in. Pdf a comparative study of fish assemblage and diversity. A tribute to claude shannon 19162001 and a plea for. A measure used by ecologists when a system contains too many individuals for each to be identified and examined.
In particular, for a random sample, we can use shannon s index of diversity aka as shannon weiner s index, which is. The shannonweiner index is most sensitive to the number of species in a sample, so it is usually considered to be biased toward measuring species richness. Shannon, simpson, and fisher diversity indices and species richness. A diversity index also called phylogenetic or simpsons diversity index is a quantitative measure that reflects how many different types such as species there are in a dataset a community and that can simultaneously take into account the phylogenetic relations among the individuals distributed among those types, such as richness, divergence or evenness. Like simpsons index, shannon s index accounts for both abundance and evenness of the species present. Distribution, diversity and ecology of spider species at two different habitats mithali mahesh halarnkar and ik pai department of zoology, goa university, india submission. The methods on offer range from the familiar, such as the calculation of various diversity indices and the fitting of common distributions, to more recentlydeveloped techniques such as total species complement estimators. Shannonwiener index h most commonly used index of diversity in ecological studies. Statistics shannon wiener diversity index in the literature, the terms species richness and species diversity are sometimes used interchangeably. Biodiversity calculator for the simpson and shannon indexes. Species richness, diversity and abundance of some decapod crustaceans in coastal waters of ondo state, south west, nigeria.